Bio Sketch


People in the Lab


Courses Taught

Bio Sketch

B.S. 1980 (Oregon State University)
M.S. 1984 (University of Maryland)
Ph.D. 1990 (University of Maryland)

Alex Flecker is a Professor in the Department of Ecology and Evolutionary Biology. He is a member of the graduate fields of Ecology and Evolutionary Biology, International Development, Natural Resources, and Conservation and Sustainable Development, and is an Associate Member of the Latin American Studies Program. After receiving his Ph.D. in Zoology (Ecology and Evolution concentration) in 1990, he was a Postdoctoral Fellow at the University of Otago in New Zealand (1991-1993), and joined the Cornell faculty in 1995. His lab has worked extensively in the Neotropics, including research sites in Venezuela , Bolivia, Ecuador, Mexico, Peru, Trinidad, and the Dominican Republic .  In addition to teaching at Cornell, he has served as an invited course instructor in Costa Rica for the Organization for Tropical Studies and at the University of Coimbra in Portugal.



Kwon, S. Y., P. M. McIntyre, A. S. Flecker, and L. Campbell. 2012. Mercury biomagnification in the food web of a neotropical stream. Science of the Total Environment 417-418:92-97, doi:10.1016/j.scitotenv.2011.11.060.


Hall, Jr., R. O., B. W. Taylor, and A. S. Flecker. 2011. Detritivorous fish indirectly reduce insect secondary production in a tropical reiver. Ecosphere 2(12):135, doi:10.1890/ES11-00042.1.


Anderson, J.T., T. Nuttle, J.S. Saldaña Rojas, T.H. Pendergast, and A.S. Flecker. 2011. Extremely long-distance seed dispersal by an overfished Amazonian frugivore.  Proceedings of the Royal Society B. 278:3329-3335, doi: 10.1098/rspb.2011.0155.


Boyero, L., and others.  2011. A global experiment suggests climate warming will not accelerate litter decomposition in streams but might reduce carbon sequestration.  Ecology Letters 14: 289–294.


Capps, K..A., M.T. Booth, S.M. Collins, M.A. Davison, J.M. Moslemi, J.L. Simonis, R.W. El-Sabaawi, and A.S. Flecker.  2011.  Nutrient diffusing substrata: a field comparison of commonly used methods to assess nutrient limitation.  Journal of the North American Benthological Society. 30: 522-532.


Flecker, A.S., P.B. McIntyre, J.W. Moore, J.T. Anderson, B.W. Taylor, and R.O. Hall.  2010. Migratory fishes as material and process subsidies in riverine ecosystems. American Fisheries Society Symposium 73:559-592.


Bassar, R.D., M.C. Marshall, A. López-Sepulcre, E. Zandonà, S.K. Auer, J. Travis, C.M. Pringle, A.S. Flecker, S.A. Thomas, D.F. Fraser, and D.N. Reznick.  2010.  Local adaptation in Trinidadian guppies alter ecosystem processes.   Proceedings of the National Academy of Sciences 107: 3616-3621.  www.pnas.org/cgi/doi/10.1073/pnas.0908023107. 


McIntyre, P.B. and A.S. Flecker. 2010. Ecological stoichiometry as in integrative framework in stream fish ecology. American Fisheries Society Symposium 73:539-558.


Winemiller, K.O., A.S. Flecker, and D.J. Hoeinghaus.  2010.  Patch dynamics and environmental heterogeneity in lotic ecosystems.  Journal of the North American Benthological Society 29:84–99.


Corman, J.R., P.B. McIntyre, B. Kuboja, W. Mbemba, D. Fink, C.W. Wheeler, C. Gans, E. Michel, and A.S. Flecker.  2010.  Upwelling couples chemical and biological dynamics in the pelagic and littoral zones of Lake Tanganyika, East Africa.  Limnology and Oceanography 55:214–224.


Anderson, J.T., J. Saldaña Rojas, and A.S. Flecker.  2009. High quality seed dispersal by fruit-eating fishes in Amazonian floodplain habitats.  Oecologia 161:279–290.


Flecker, A.S.  2009.  Crossing borders: promoting graduate research in the developing world: Faculty Response.  Frontiers in Ecology and the Environment 7: 334.


Knoll, L.B., P.B.McIntyre, M.J. Vanni, and A.S. Flecker. 2009.  Feedbacks of consumer nutrient recycling on producer biomass and stoichiometry: separating direct and indirect effects.  Oikos 118:1732-1742.


McIntyre, P.B., A.S. Flecker, M.J. Vanni, J.M. Hood, B.W. Taylor, and S.A. Thomas.  2008.  Fish distributions and nutrient recycling in a Neotropical stream: can fish create biogeochemical hotspots?  Ecology 89: 2335–2346.


Power, A.G. and A.S. Flecker.  2008.  The role of vector diversity in disease dynamics.  Pp.30-47 In: Ostfeld, R. S., F. Keesing, and V. Eviner (eds.).  Infectious Disease Ecology: Effects of Disease on Ecosystems and of Ecosystems on Disease.  Princeton University Press, Princeton, NJ.


McIntyre, P.B., L.E. Jones, A.S. Flecker, and M.J. Vanni.   2007.  Fish extinctions alter nutrient recycling in tropical freshwaters.  Proceedings of the National Academy of Sciences 104: 4461-4466.


Taylor, B.W., A.S. Flecker, and R.O. Hall, Jr. 2006.  Loss of a harvested fish species disrupts carbon flow in a diverse tropical river.  Science 313:833-836. pdf


McIntyre, P.B. and A.S. Flecker.  2006.   Rapid turnover of tissue nitrogen of primary consumers in tropical freshwaters.  Oecologia 148: 12–21.  pdf


Allan, J.D., A.S. Flecker, S. Segnini, D.C. Taphorn, E. Sokal and G.W. Kling.  2006.   Limnology of Andean piedmont rivers of Venezuela.  Journal of the North American Benthological Society 25: 66–81.  pdf


Hood, J.M., M.J. Vanni, and A.S. Flecker.  2005.  Nutrient recycling by two phosphorus-rich grazing catfish: the potential for phosphorous limitation of fish growth.  Oecologia 146: 247–257. pdf


Templer, P.H., P.M. Groffman, A.S. Flecker, and A.G. Power.  2005.  Land use change and soil nutrient transformations in the Los Haitises region of the Dominican Republic.  Soil Biology & Biochemistry 37: 215–225.  pdf

Solomon, C.T., A.S. Flecker, and B.W. Taylor.  2004. Sediment-mediated interactions between tadpoles and armored catfish in a Neotropical stream.  Copeia 2004:610-616.  pdf

Uriarte, M., L. W.  Rivera, J. K. Zimmerman, T. M. Aide, A.G. Power and A. Flecker.  2004.  Interactions between land use history and hurricane damage in a Neotropical forest.  Plant Ecology 174: 49-58.  pdf

Flecker, A.S. and B.W. Taylor.  2004.  Tropical fishes as biological bulldozers: density effects on resource heterogeneity and species diversity. Ecology 85(8):2267-2278. pdf

McIntyre, P.B., S. Baldwin, and A.S. Flecker.  2004.  Effects of behavioral and morphological plasticity on risk of predation in a Neotropical tadpole.  Oecologia 141:130-138. pdf

Wright, J.P. and A.S. Flecker. 2004. Deforesting the riverscape: The effects of wood removal on fish diversity in a Venezuelan piedmont stream. Biological Conservation 120:443-451. pdf

Riley, R.H., C.R. Townsend,  D.A. Raffaelli, and A.S. Flecker.   2004.  Sources and effects of subsidies along the stream-estuary continuum. pp. 241-260  In: G.A. Polis, M.E. Power, and G.R. Huxel (eds.).  Food webs at the landscape level.  University of Chicago Press, Chicago, IL

Wright, J.P., A.S. Flecker, and C.G. Jones.  2003.  Local vs. landscape controls on plant species richness in beaver meadows.  Ecology 84:3162-3173. pdf

Power, A.G. and A.S. Flecker.  2003.  Virus specificity in disease systems: Are species redundant?  pp. 330-346.  In: P. Kareiva and S. A. Levin (eds.). The importance of species: perspectives on expendability and triage. Princeton University Press, Princeton, NJ pdf

Flecker, A.S., B.W. Taylor, E.S. Bernhardt, J.M. Hood, W.K. Cornwell, S.R. Cassatt, M.J. Vanni, and N.S. Altman. 2002. Interactions between herbivorous fishes and limiting nutrients in a tropical stream ecosystem. Ecology 83:1831-1844. pdf

Vanni, M.J., A.S. Flecker, J.M. Hood, and J.L. Headworth. 2002. Stoichiometry of nutrient recycling by vertebrates in a tropical stream: linking species identity and ecosystem processes. Ecology Letters 5: 285-293 pdf

Wright, J.P., C.G. Jones, and A.S. Flecker. 2002. An ecosystem engineer, the beaver, increases species richness at the landscape scale. Oecologia 132: 96-101 pdf

Flecker, A.S. 1997. Habitat modification by tropical fishes: environmental heterogeneity and the variability of interaction strength. Journal of the North American Benthological Society 16:286-295

Flecker, A.S. 1996. Ecosystem engineering by a dominant detritivore in a diverse tropical ecosystem. Ecology 77: 1845-1854. pdf

Flecker, A. S. and C. R. Townsend. 1994. Community-wide consequences of trout introduction in New Zealand streams. Ecological Applications 4:798-807

Flecker, A. S. and B. P. Feifarek. 1994. Disturbance and the temporal variability of invertebrate assemblages in two Andean streams. Freshwater Biology 31:131-142

Allan, J. D. and A. S. Flecker. 1993. Biodiversity conservation in running waters. BioScience 43:32-43. pdf

Flecker, A. S. 1992. Fish trophic guilds and the structure of a tropical stream: Weak direct versus strong indirect effects. Ecology 73:927-940. pdf

Flecker, A. S. 1992. Fish predation and the evolution of invertebrate drift periodicity: Evidence from neotropical streams. Ecology 73:438-448 pdf